Reply to Risk and Zhu: Mixed-effects modeling as a principled approach to heritability analysis with repeat measurements. Ge T, Holmes AJ, Buckner RL, Smoller JW, Sabuncu MR. Reply to Risk and Zhu: Mixed-effects modeling as a principled approach to heritability analysis with repeat measurements. Proc Natl Acad Sci U S A. 2018;115(2):E123. doi:10.1073/pnas.1719882115 Read more